Hugo de Vries's contribution is

Options :
  1. theory of natural selection
  2. theory of mutations
  3. law of dominance
  4. law of segregation
Answer and Explanation :-

Answer: Option 2

Explanation:

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For the future, I shall rely only upon those elements of my character which I have tested. Who would ever have said that I should find pleasure in shedding tears? That I should love the man who proves to me that I am nothing more than a fool?
-Stendhal The Red and the Black